I am a palaeobiologist interested in the large-scale patterns of macroevolution, particularly trait evolution. My research has spanned a large part of the (eukaryotic) tree of life: I have investigated evolutionary patterns in mammals, birds, and plants. My continuing research focus is been to join fossil data to information from living species to gain a more complete understanding of the diversification of life. In addition I have explored aspects of divergence time estimation, and have evaluated the best methods to infer topology from morphological cladistic matrices.


  • 2017-Present. 1851 Research Fellow, University of Bath
  • 2016-2017. Postdoctoral researcher, Natural History Museum and University of Bristol.
  • 2012-2016. PhD “Using phylogenies to explore macroevolution” supervised by Professor Mike Benton (University of Bristol) and Dr Gavin Thomas (University of Bristol). Awarded the 2016 best PhD thesis’ award for the Faculty of Science.
  • 2011-2012. MSc Palaeobiology, Distinction. University of Bristol. Awarded the David Dineley Prize.
  • 2006-2009. BSc Biological Sciences, 1st Class Honours. University of Exeter.